BKE_mesh_new_nomain automatically added a CD_ORIGINDEX layer initialized to 0, which was never filled in correctly. In 2.7 the equivalent function used to modify the source derivedmesh and add valid original indices to it, but this is no longer possible in the new design and was quite unpredictable anyway. Now instead rely on mesh_calc_modifiers and the depsgraph to determine when CD_ORIGINDEX should be added.